Helmus Publications


  • Behm, JE, BR Waite, ST Hsieh, MR Helmus2018. Benefits and limitations of three-dimensional printing technology for ecological research. bioRxiv 283895. 
  • Marin, J, G Rapacciuolo, GC Costa, CH Graham, TM Brooks, BE Young, VC Radeloff, JE BehmMR HelmusSB Hedges. 2018. Evolutionary time drives global tetrapod diversity. Proceedings of the Royal Society B 285 (1872), 1-8. 
  • Helmus, MR and JE Behm. 2018. Anthropocene Island Biogeography. In: Encyclopedia of the Anthropocene. (eds. D. DellaSala, M. Goldstein). Elsevier. pp 37-43. Book chapter.


  • Rapacciuolo, G, J Marin, GC Costa, MR Helmus, JE Behm, TM Brooks, SB Hedges, VC Radeloff, BE Young, CH Graham. 2017. The signature of human pressure history on the biogeography of body mass in tetrapods. Global Ecology and Biogeography 1-13.
  • Tucker, CM, MW Cadotte, SB Carvalho, TJ Davies, S Ferrier, SA Fritz, R Grenyer, MR Helmus, LS Jin, AO Mooers, S Pavoine, O Purschke, DW Redding, DF Rosauer, M Winter, F Mazel. 2017. A guide to phylogenetic metrics for conservation, community ecology and macroecology. Biological Reviews 92 (2), 698-715.  
  • Helmus, MR, JE Behm, WAM Jesse, JJ Kolbe, J Ellers, JB Losos. 2017. Exotics exhibit more evolutionary history than natives: A comparison of the ecology and evolution of exotic and native anole lizards. Invasion Genetics: The Baker and Stebbins Legacy, 122-138. 


  • Nelson, EJ, MR Helmus, J Cavender-Bares, S Polasky, JR Lasky, AE Zanne, WD Pearse, NJB Kraft, DA Miteva, WF Fagan. 2016. Commercial Plant Production and Consumption Still Follow the Latitudinal Gradient in Species Diversity despite Economic Globalization. PloS One 11 (10), e0163002.  
  • Yguel, B, H Jactel, IS Pearse, D Moen, M Winter, J Hortal, MR Helmus, I Kühn, S Pavoine, O Purschke, E Weiher, C Violle, W Ozinga, M Brändle, I Bartish, A Prinzing. 2016. The evolutionary legacy of diversification predicts ecosystem function. The American Naturalist 188 (4), 398-410. 
  • Behm, JE and MR Helmus. 2016. Microhyla butleri (Tubercled Pygmy frog) predation. Herpetological Review, 47(4): 643-644. 


  • Pearse, WD, MW Cadotte, J Cavender-Bares, AR Ives, CM Tucker, SC Walker, MR Helmus. 2015. pez: phylogenetics for the environmental sciences. Bioinformatics 31 (17), 2888-2890. 
  • Vermaat, JE, J Ellers, MR Helmus. 2015. The role of biodiversity in the provision of ecosystem services. Ecosystem Services: From Concept to Practice, 25. Email for pdf.


  • Helmus, MR, DL Mahler, JB Losos. 2014. Island biogeography of the Anthropocene. Nature 513 (7519), 543-546.  
  • Vasseur, DA, JW Fox, A Gonzalez, R Adrian, BE Beisner, MR Helmus, C Johnson, P Kratina, C Kremer, C de Mazancourt, E Miller, WA Nelson, M Paterson, JA Rusak, JB Shurin, CF Steiner. 2014. Synchronous dynamics of zooplankton competitors prevail in temperate lake ecosystems. Proc. R. Soc. B 281 (1788).  
  • Pearse, WD, A Purvis, J Cavender-Bares, MR Helmus. 2014. Metrics and Models of Community Phylogenetics. Modern Phylogenetic Comparative Methods and Their Application in Evolutionary Biology. Springer Berlin Heidelberg, pp. 451-464. 


  • Blanchet S, MR Helmus, S Brosse, G Grenouillet. 2013. Regional vs local drivers of phylogenetic and species diversity in stream fish communities. Freshwater Biology 59 (3), 450-462. 
  • Helmus, MR, N Mercado-Silva, MJ Vander Zanden. 2013. Subsidies to predators, apparent competition and the phylogenetic structure of prey communities. Oecologia 173 (3), 997-1007. 


  • Kua, CS, J Ruan, J Harting, CX Ye, MR Helmus, J Yu, CH Cannon. 2012. Reference-Free Comparative Genomics of 174 Chloroplasts. PloS One 7 (11), e48995. 
  • Helmus, MR, AR Ives. 2012. Phylogenetic diversity–area curves. Ecology 93:S1-S43.  


  • Eklöf, A, MR Helmus, M Moore, S Allesina. 2011. Relevance of evolutionary history for food web structure. Proceedings of the Royal Society of London B: Biological Sciences 279 (1733), 1588-1596. 
  • Ives, AR, MR Helmus. 2011. Generalized linear mixed models for phylogenetic analyses of community structure. Ecological Monographs 81 (3), 511-525.  
  • Scheiner, SM, A Chiarucci, GA Fox, MR Helmus, DJ McGlinn, MR Willig. 2011. The underpinnings of the relationship of species richness with space and time. Ecological Monographs 81 (2), 195-213. 

2010 and earlier

  • Ives, AR, MR Helmus. 2010. Phylogenetic metrics of community similarity. The American Naturalist 176, E128-E142. 
  • Kembel, SW, PD Cowan, MR Helmus, WK Cornwell, H Morlon, DD Ackerly, SP Blomberg, CO Webb. 2010. Picante: R tools for integrating phylogenies and ecology. Bioinformatics 26 (11), 1463. 
  • Helmus, MR, WB Keller, MJ Paterson, ND Yan, CH Cannon, JA Rusak. 2010. Communities contain closely related species during ecosystem disturbance. Ecology Letters 13 (2), 162-174. 
  • Ahrenstorff, TD, GG Sass, MR Helmus. 2009. The influence of littoral zone coarse woody habitat on home range size, spatial distribution, and feeding ecology of largemouth bass (Micropterus salmoides). Hydrobiologia 623 (1), 223-233. 
  • Helmus, MR, LB Allen, O Dominguez-Dominguez, ED Pardo, P Gesundheit, J Lyons, N Mercado-Silva. 2009. Threatened fishes of the world: Allotoca goslinei Smith and Miller, 1987 (Goodeidae). Environmental Biology of Fishes 84 (2), 197-198. 
  • Mercado‐Silva, N, MR Helmus, M Zanden. 2008. The effects of impoundment and non‐native species on a river food web in Mexico’s central plateau. River Research and Applications 25 (9), 1090-1108. 
  • Helmus, MR, GG Sass. 2008. The rapid effects of a whole‐lake reduction of coarse woody debris on fish and benthic macroinvertebrates. Freshwater Biology 53 (7), 1423-1433. 
  • Newton, RJ, SE Jones, MR Helmus, KD McMahon. 2007. Phylogenetic ecology of the freshwater Actinobacteria acI lineage. Applied and Environmental Microbiology 73 (22), 7169-7176. 
  • Helmus, MR, K Savage, MW Diebel, JT Maxted, AR Ives. 2007. Separating the determinants of phylogenetic community structure. Ecology Letters 10 (10), 917-925.  
  • Helmus, MR, TJ Bland, CK Williams, AR Ives. 2007. Phylogenetic measures of biodiversity. The American Naturalist 169 (3), E68-E83. 
  • Helmus, MR, DE Dussourd. 2005. Glues or poisons: which triggers vein cutting by monarch caterpillars? Chemoecology 15 (1), 45-49. 

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